Full statistics and more than you will probably ever need are built right into the site. Clicks, pageviews, time spent on site, which feeds are used and how often, most, e-mails, most viewed, etc. Ads can be tracked through sent e-mails, but amount of times read through e-mail (automatic e-mails like news alerts) cannot be tracked. This is because once the story is e-mailed, it leaves the server and cannot be tracked. However, the ads associated with the automatic e-mails will reside on the server, and therefore can still be tracked for statistical purposes.
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Google Analytics
Google Analytics is an awesome (free) program from Google which we recommend as well. More info here: http://www.google.com/analytics